Agrotis gravis Grote, 1874
93-3518
WA : Island Co.
Deception Pass SP, South Beach, 23 ft
48.4, -122.7
May 06, 1995, L. G. Crabo.
Specimen courtesy of LGCC
Photograph copyright: Merrill A. Peterson
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Identification
Adults
Agrotis gravis is a distinctive, small to medium-sized (FW length 13 - 16 mm), light brown-gray to blackish-brown Agrotis that flies on Pacific Coast beaches in late spring and early summer. It has an oblique orbicular spot and a large reniform spot that is drawn toward the forewing base at the posterior end. The forewing ground color is variable, ranging from mottled, medium dark blue-gray, brown-gray, gray-brown, dull red-brown, or dark blackish-brown. Lighter colored specimens have a darker costa, often with a red-brown patch between the postmedial and subterminal lines, and dark gray to black margin. The transverse lines are black and double, but relatively indistinct except near the costa. The antemedial line is irregular and the postmedial line is dentate. The pale subterminal line is extended to the margin on veins M3 and CuA1, forming a pale "W" against the dark margin, and is preceded by black wedges in the middle of the forewing. The orbicular and reniform spots are black with an inner pale ochre or gray line and dark central filling. The orbicular spot is oval, occasionally fused to the reniform spot. The reniform spot is large and drawn toward the base along the cubital vein in most specimens. The light inner component of the medial part of the spot is the most conspicuous part. The claviform spot is black and moderately large. The hindwing is pale gray with a slight brownish tint basally and with a smoky gray submarginal to marginal band, discal spot, and veins, and a gray fringe in males; it is entirely darker gray in females. The ground color of the thorax matches the forewing. The collar is striped with transverse black lines and a light tip. The tegulae are light gray to brown with darker gray margins. The male antenna is weakly bipectinate.
This species is only likely to be confused with Agrotis vancouverensis, a widespread and common Agrotis that occurs in the western portion Pacific Northwest, including on beaches. This species is usually warmer brick-red than A. gravis, but it too is variable in color and there is overlap of gray and brown color between these species. The forewing shape of A. vancouverensis is less pointed that that of A. gravis, most specimens of A. vancouverensis lack the long chevron-shaped black dashes before the subterminal line (these spots are shorter and lack a lateral notch if present in A. vancouverensis), the orbicular spot of A. vancouverensis is rounder and lacks the prominent pale inner line of A. gravis, and the reniform spot is smaller and kidney-shaped without the extension toward the base that is common in A. gravis.
Habitat
This species is narrowly endemic to sand dunes adjacent to the Pacific Ocean that are covered with beach grass and coastal herbaceous vegetation, or coastal grasslands with a sand-humus substrate. It is usually common to abundant in this very restricted habitat.
Distribution
Pacific Northwest
Agrotis gravis is found on the immediate Pacific Coast as far north as south-western British Columbia.
Global
The northern limits of this species is reached in our region. Further south it occurs in coastal California north of the Transverse Ranges.
Life History
Larvae
This species is a sand-burrowing cutworm that feeds on general herbaceous vegetation.
Adults
The species is single-brooded with a long flight period that lasts from early April to early July. It is nocturnal and comes to lights.
Economic Importance
None.
Literature
Crumb (1956)
Lafontaine (2004)