Euxoa pallidimacula Lafontaine, 1987
Euxoa pallidimacula is a medium-size Euxoa (FW length 14–18 mm) with, as the name suggests, a pale gray-tan forewing with a dark median line and pale filling of the orbicular and reniform spots that flies in dry steppe during the early summer. The ground color varies slightly from pallid to medium-dark gray-tan, lightly suffused with dark scales. The dominant dark mark is the gray median line, but the terminal and distal subterminal areas are also darker gray. The basal and antemedial lines are similar, dark gray, double with components of unequal strength filled with the ground color. The antmedial line has darker lateral and weaker medial components and is slightly irregular and angled slightly toward the outer margin. The median line is dark gray, prominent, moderately diffuse and moderately wide, slightly angled below the reniform spot where there is a focal concave defect in the medial edge. The postmedial line is dark, toothed on the veins, usually double with a light gray outer component, drawn basad on the costa, smoothly convex or laterally blunted around the reniform spot, and angled slightly toward the base below the level of the cell. The subterminal line is pale tan, slightly irregular with a moderate W-mark on veins M3 and CuA1. It is preceded by a dark gray shade, strongest on and near the costa. The terminal line is dark gray, thin or a series of spots. The fringe is similar to the terminal area. The orbicular and reniform spots often lack dark outlines—the orbicular spot is partially lined with dark gray in a few specimens— and are filled with a pale gray-tan. The orbicular spot is oval, fairly small. The reniform spot is kidney-shaped, moderate-sized but fading into the adjacent median area laterally. No claviform spot is present. The hindwing is tan-gray with slighly darker gray margin, discal spot, and terminal line. The hindwing fringe is white with a tan base. The head and thorax match the forewing color and lack dark marks. A frontal tubercle is present. The male antenna is biserrate with long fasciculae. Euxoa pallidimacula is a member of the subgenus Euxoa.
This species can be recognized by its early summer flight, pallid forewing color with gray lines dominated by the median line, and relatively inconspicuous pale-filled forewing spots. It is very close in appearance to Euxoa setonia, a more widespread and common species that usually flies in forests. They differ mainly in the darkness and strength of the markings, with E. setonia being darker tan with dark transverse lines and often with yellowish filling of the spots. Males of these species differ structurally in the orientation of the saccular extension of the valve, which is visible after brushing the scales from the end of the abdomen. Those of E. pallidimacula are straight and are parallel to the ventral valve margin, while those of E. setonia are curved slightly ventrad away from the valve. Other light gray-tan early-season Euxoa species with which E. pallidimacula fly are either less strongly marked with dark, such as Euxoa scotogrammoides, or have stronger dark markings with dark outlines of the spots. An example of the latter is Euxoa terrenus, a dark brown species in forests that is paler in arid habitats.
Euxoa is a very large genus with many similar species, many of which are also quite variable. The genus is defined by a saccular extension of the valves in males and sclerotized plates on the dorsal and ventral ductus bursae in females. The genus was revised by Lafontaine (1987) in the Moths of North America series and is divided into eight subgenera based on structural characters. These moths are amongst the most difficult to identify. Even though the forewing color and strength of various markings can vary significantly in some of the species, the shape of the lines and spots and the color of the hindwings (often different in the sexes) are more constant. The habitat and flight period are also important in helping to narrow the possibilities.
This species is narrowly endemic to dry rangeland habitats. In the Pacific Northwest, it is found sporadically in dry sagebrush steppe and open juniper woodland habitats.
Euxoa pallidimacula is found in the western portion of the Columbia Basin and in southeastern Oregon. This species was first collected in the Pacific Northwest after the original species description and are therefore not included in the description of the range in the MONA fascicle (Lafontaine 1987)
This species is found in the Great Basin and on the Colorado Plateau to the south of the Pacific Northwest. The range extends west to eastern California and south to northern Arizona.
No information is presently available regarding larval foodplants of this species, but it is probably a soil-surface feeding cutworm that feeds on herbaceous vegetation based on related species.
Euxoa pallidimacula flies in the late spring and early summer. Most of the records from our region are from late May and June. It is nocturnal and comes to lights.