Euxoa tocoyae (Smith, 1900)
Euxoa tocoyae is an orange-brown medium-size Euxoa (FW length 16–19 mm) with a faint pattern lacking strongly contrasting dark gray or black elements that flies in dry forests south of the central Washington Cascades during late spring and early summer. Larger specimens of this moth are relatively stocky for the genus. The forewing is medium-dark gray-tan, strongly tinted orange or red-orange. The gray-brown median line and shade preceding the subterminal line are usually darkest, although the terminal area often contains gray, but are usually not strongly contrasting. The basal, antemedial, and postmedial lines are gray-brown, double filled with slightly lighter shade of the ground color, darkest on the costa and variable in darkness across the rest of the wing. The basal line is inconspicuous or evident as spots on the costa. The antemedial line is usually present across the wing, occasionally obsolete below the costa, slightly irregular and angled slightly toward the outer margin. The median line is gray-brown, irregular, coursing through the reniform spot and bent apex lateral below the spot. The postmedial line is toothed weakly on the veins when evident, bluntly excurved around the reniform spot and angled toward the base below the level of the cell. The subterminal line is pale, indistinct and often patchy, irregular with an offset below the costa and a moderate W-mark on the mid-wing on veins M3 and CuA1. It is preceded by an even gray-brown shade. The terminal line is thin or broken into spots. The fringe matches the terminal area. The orbicular and reniform spots are outlined in a pale shade of the ground and are relatively small and inconspicuous, sometimes absent in poorly marked specimens. The orbicular spot is round, filled with the ground color. The reniform spot is kidney-shaped, filled with the ground color and a darker gray spot in the lower end, occasionally entirely darker gray. The claviform spot is absent. The hindwing is light brownish gray, with slightly darker veins, discal spot, and terminal line. The hindwing fringe is white with ochre and gray base. The head and thorax match the forewing, sometimes with a very weak and thin gray line across the mid-collar. A frontal tubercle is present. The male antenna is weakly bipectinate. Euxoa tocoyae is a member of the large subgenus Euxoa.
This species is fairly non-descript in pattern but can usually be recognized by its relatively early flight period, the orange-tan color, pale-rimmed spots, and the lack of strongly contrasting lines. It is similar to Euxoa scotogrammoides, its closest relative in our area, but it is gray-brown and has a smaller reniform spot that is more uniformly filled with dark than that of E. tocoyae. Euxoa comosa, a common and variable species that usually flies slightly later in the season than E. tocoyae has orange and reddish forms. These usually have darker lines than E. tocoyae, and its spots are at least partially outlined by thin dark lines. Euxoa nostra is an orange-brown species that also flies during the mid-summer. Most specimens have a powdery gray suffusion along the costa that E. tocoyae lacks. The few specimens lacking this suffusion differ in having dark outlines around the spots, and an oval rather than round orbicular spot.
Euxoa is a very large genus with many similar species, many of which are also quite variable. The genus is defined by a saccular extension of the valves in males and sclerotized plates on the dorsal and ventral ductus bursae in females. The genus was revised by Lafontaine (1987) in the Moths of North America series and is divided into eight subgenera based on structural characters. These moths are amongst the most difficult to identify. Even though the forewing color and strength of various markings can vary significantly in some of the species, the shape of the lines and spots and the color of the hindwings (often different in the sexes) are more constant. The habitat and flight period are also important in helping to narrow the possibilities.
This species flies in dry forest and grassland habitats. It is rare and sporadic east of the Cascades, but is common in dry grasslands, oak savanna, oak woodlands, and mixed hardwood-conifer forests at low to middle elevations west of the Cascades, particularly in southwestern Oregon. It flies in open mixed forest in southern Washington and in riparian forest near steppe in southeastern Oregon.
Euxoa tocoyae is found from the east slope of the Cascades in southern Washington through eastern and southwestern Oregon. It has been most commonly collected in the Siskiyou Mountains of southwestern Oregon in our region. The most easterly occurrence is at the base of Steens Mountain in Harney County, Oregon.
This moth is found near the Pacific Coast of North America. The range extends south through California to San Diego County near the border with Mexico.
No information is presently available regarding larval foodplants of this species, but it is probably a soil-surface feeding cutworm that feeds on herbaceous vegetation based on related species.
This species is usually found in late spring and early summer. The records from our region extend from mid-May to late June.